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Eyen, Khoo and Tsen, Sandy Tze Lui and Lee, Yew Leung and John Baptist Sugau and Joan Teresa Pereira and Reuben Nilus and Maycock, Colin Ruzelion (2022) Upuna Symington. In: The Sabah Plant Red Listing of Dipterocarpaceae. Jabatan Perhutanan Sabah, Sandakan, Sabah, Malaysia, pp. 186-187. ISBN 978-967-0180342
